Open research questions in Fossil Insects in Amber
70 gap statements mined from Fossil Insects in Amber papers in our 4.5M-paper local library, which holds 304 papers on the topic — drawn mostly from each paper's own stated research gap, future-work, challenge and limitation notes, and its abstract. The ones listed below are a selection still marked open; each names the study that raised it, with a DOI link where the paper has one.
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What the literature leaves open
Further study of the fossil record of Evanioidea is needed to understand the evolution of the group. The discovery of A. hirsuta highlights the importance of continued exploration of Cretaceous amber deposits.
A new lineage of evaniiform wasps in Cretaceous amber from Myanmar (Hymenoptera: Evanioidea) · 2026 · DOIThe larval stage of most species of the genus Allopodagrion is unknown. The relationship between the larva and the adult of A. contortum was not previously confirmed.
Integrative taxonomy reveals the larva of Allopodagrion contortum (Hagen in Selys, 1862) (Odonata: Megapodagrionidae) from Southeastern Brazil · 2026 · DOIFurther study of the Jiyuan Basin fossil site is needed. More research on the biostratigraphic correlation between the Daohugou and Jiyuan fossil localities is required.
A new species of Proptychopterina (Diptera: Nematocera: Ptychopteridae) from the Middle Jurassic of the Jiyuan Basin, China · 2026 · DOIThere is a lack of description of insect fossils from the Jiyuan Basin. The biostratigraphic correlation between the Daohugou and Jiyuan fossil localities was unclear.
A new species of Proptychopterina (Diptera: Nematocera: Ptychopteridae) from the Middle Jurassic of the Jiyuan Basin, China · 2026 · DOIFurther study is needed to fully understand the characteristics of the amber from the Jiufengshan Formation. Additional research is required to reconstruct the palaeovegetation of the Jiufengshan Formation. More work is needed to develop new methods for analyzing amber and other fossilized materials.
Early Cretaceous amber from the Heibaoshan-Handaqi Basin, Heilongjiang Province, Northeast China · 2026 · DOIFurther study of the Kachin amber beetle fauna is needed to fully realize its potential for understanding beetle evolution and diversity. The use of modern imaging techniques, such as confocal microscopy and micro-computed tomography, could be further developed and applied to the study of fossil specimens.
There is a lack of understanding of the unique characters of the mid-Cretaceous genus Punctocorydasialis. There is a need for more research on the evolution of Neuropterida.
New unique characters of the mid-Cretaceous genus Punctocorydasialis Chen et al., 2026 (Neuropterida: Corydasialidae) · 2026 · DOIThe phylogenetic placement of the genera Euphilotes and Philotiella remains uncertain. In particular, the limits, composition, and diagnostic characters of several lineages remain insufficiently resolved and will require further phylogenetic, morphological, and biological study before robust diagnoses can be established.
The study of fossil material can be challenging due to the limited preservation of anatomical structures. The delimitation of fossil taxa can be difficult due to the lack of complete specimens. The comparison of fossil and extant species can be challenging due to the differences in preservation and morphology.
Hidden but unearthed: an unusual new genus and three new species for the planthopper family Nogodinidae (Hemiptera: Fulgoromorpha) from Early Miocene Mexican amber, with a key to New World Nogodinini · 2026 · DOIThe taxonomy of Devonian trilobites from the Cantabrian Mountains has received little attention since the principal contributions by earlier researchers. A systematic revision is necessary to bring the outdated nomenclature into line with contemporary frameworks.
Revised and new Emsian (Lower Devonian) trilobites from Polentinos, Cantabrian Mountains, northern Spain · 2026 · DOIThe paper mentions the need for controlled conditions during resin extraction to prevent clogging of the resin canals.
Resin does not safeguard insect DNA during early decay despite preservation of soft tissues · 2026 · DOIThe fossil is preserved in a piece of amber with uneven surfaces, which slightly impacts some views. Further trimming and polishing of the amber was impossible due to the closeness of the wasp to the amber surface.
A new lineage of evaniiform wasps in Cretaceous amber from Myanmar (Hymenoptera: Evanioidea) · 2026 · DOIThe molecular diversity of the genus Ecdyonurus is high, making it challenging to resolve taxonomic relationships. The study of fossil mayflies can be challenging due to the limited availability of specimens. The use of COI barcoding and phylogenetic analysis can be limited by the quality of the sequences and the choice of methods.
A new mayfly species of Ecdyonurus Eaton, 1868 from Eocene Baltic Amber and insights into the extant molecular diversity of the genus · 2026 · DOINew and little known species of Sphecidae of Eastern Siberia and Far East of USSR.
A revision of the Palearctic species of the genus Crabro Fabricius, 1775 (Hymenoptera: Spheciformes: Crabronidae), with reclassification of the world subgenera · 2026 · DOIThe study seeks to provide new data for studies of Mesozoic amber in China.
Early Cretaceous amber from the Heibaoshan-Handaqi Basin, Heilongjiang Province, Northeast China · 2026 · DOIThe study of fossil specimens can be challenging due to the limited availability of material and the difficulty of interpreting fossil evidence. The use of modern imaging techniques requires specialized expertise and equipment.
Prior studies have addressed higher-level relationships within the Saturniidae, but with sparse taxon and often reduced gene sampling.
Phylogenomics Illuminates the Evolutionary History of Wild Silkmoths in Space and Time (Lepidoptera: Saturniidae) · 2026 · DOIThe study only examined two subfamilies of Aphalaridae. The molecular dataset was limited to 24 species. The study did not investigate the functional significance of cuticular microsculpture.
Beyond the obvious: diversity and evolution of cuticular microsculpture in jumping plant lice of the subfamilies Aphalarinae and Rhinocolinae (Hemiptera, Psylloidea, Aphalaridae) · 2026 · DOIImproved species identification in genetic databases - Development of alternative molecular markers - Expansion of taxon sampling to resolve taxonomic…
A new mayfly species of Ecdyonurus Eaton, 1868 from Eocene Baltic Amber and insights into the extant molecular diversity of the genus · 2026 · DOIAccounting for sampling bias in fossil insect assemblages. Propagating uncertainty through successive trophic steps. Integrating independent geochemical indicators and other plant data methods to cross-validate and further calibrate the reconstruction approach.
Reconstructing vegetation biomass in the Middle Jurassic Yanliao Biota from insect fossil assemblages · 2026 · DOIFuture research should integrate independent geochemical indicators and other plant data methods to cross-validate and further calibrate this reconstruction approach, - Research should focus on comparative analyses of terrestrial ecosystems through deep…
Reconstructing vegetation biomass in the Middle Jurassic Yanliao Biota from insect fossil assemblages · 2026 · DOIIt is not difficult to imagine that large stag beetles are difficult to preserve in amber, and since the modern Lucaninae have a high proportion of large species, some caution is warranted in this regard…
Taxonomic notes on Aesalinae stag beetles (Coleoptera: Lucanidae), with special reference to those in mid-Cretaceous Kachin amber · 2026 · DOIFurther investigation of the functional significance of cuticular microsculpture. Examination of cuticular microsculpture in other subfamilies of Aphalaridae. Investigation of the evolution and diversity of cuticular microsculpture in other insect groups.
Beyond the obvious: diversity and evolution of cuticular microsculpture in jumping plant lice of the subfamilies Aphalarinae and Rhinocolinae (Hemiptera, Psylloidea, Aphalaridae) · 2026 · DOIFurther study of fossil material to better understand the evolution and diversity of Nogodinidae. Phylogenetic analysis of the new genus and species to determine their relationships to other members of the tribe. Investigation of the biology and ecology of planthoppers to inform conservation efforts and ecosystem management.
Hidden but unearthed: an unusual new genus and three new species for the planthopper family Nogodinidae (Hemiptera: Fulgoromorpha) from Early Miocene Mexican amber, with a key to New World Nogodinini · 2026 · DOIInvestigate the preservation of insect DNA in resin over longer time periods, - Examine the effects of different types of resin on insect DNA preservation, - Study the preservation of soft tissues in other groups…
Resin does not safeguard insect DNA during early decay despite preservation of soft tissues · 2026 · DOI
Most-cited papers in Fossil Insects in Amber
- Phylogenomics resolves the timing and pattern of insect evolution · Science · 2014 · 2,460 citations
- Comparative gene expression supports the origin of the incisor and molar process from a single endite in the mandible of the red flour beetle Tribolium castaneum · EvoDevo · 2013 · 41 citations
- What genes and chromosomes say about the origin and evolution of insects and other arthropods · Russian Journal of Genetics · 2010 · 40 citations
- Phylogeography of the Coccus scale insects inhabiting myrmecophytic Macaranga plants in Southeast Asia · Population Ecology · 2009 · 35 citations
- Revision of the Tenebrionidae of America, North of Mexico · Transactions of the American Philosophical Society · 1870 · 35 citations
- North American Plesiosaurs Trinacromerum · The Journal of Geology · 1908 · 34 citations
- The Giant Resin Bee, Megachile sculpturalis Smith: New Distributional Records for the Mid- and Gulf-south USA · Biodiversity Data Journal · 2015 · 31 citations
- Descriptions of New North American Insects, and Observations on Some Already Described · Transactions of the American Philosophical Society · 1834 · 25 citations
- Rapid growth and the evolution of complete metamorphosis in insects · Proceedings of the National Academy of Sciences · 2024 · 25 citations
- The radiation of Hymenoptera illuminated by Bayesian inferences from the fossil record · Current Biology · 2025 · 23 citations
Most recent work
- Annotated checklist of Recent Polyplacophora with notes on their systematics · Zootaxa · 2026
- From setae to scales: Burmese amber fossils reveal the Early Cretaceous innovation of body coverings in scaly crickets · Proceedings of the Royal Society B Biological Sciences · 2026
- Phylogenomics Illuminates the Evolutionary History of Wild Silkmoths in Space and Time (Lepidoptera: Saturniidae) · Systematic Biology · 2026
- The rise and fall of the Eocene fly genus Kelneria (Diptera, Keroplatidae) · Arthropod Systematics & Phylogeny · 2026
- New Mammutidae material from Yushe Basin (southeastern Shanxi, China) and their implications for Mammut evolution · Palaeoworld · 2026
- Phylogenomics of Manticorini tiger beetles supports ancient Gondwanan vicariance and recent American amphitropical disjunction · Proceedings of the Royal Society B Biological Sciences · 2026
- Fossil gaps, ghost lineages, and ‘major extinction events’ · Current Biology · 2026
- A new gall midge fly (Diptera: Cecidomyiidae) from the Lowermost Eocene Oise amber (France) · Historical Biology · 2026
- A new lineage of evaniiform wasps in Cretaceous amber from Myanmar (Hymenoptera: Evanioidea) · Mesozoic · 2026
- Descriptions and biological data on three new species of Trigonalyidae (Hymenoptera: Trigonalyoidea) from South Korea · European Journal of Taxonomy · 2026
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